He has got a preliminary and upright postprotocrista labially created. The fresh preprotocrista, seemingly straight, satisfies a highly-build parastyle. The fresh new metaconule was some smaller which can be larger on the M3. The newest premetaconulecrista in addition to fossae are simple in place of neocrista. The posterior fossa is actually angled, specifically toward M3. Out of straight into the M1, the fresh postmetaconulecrista gets sigmoidal into the M3, bypassing the latest metacone. The fresh new labial cusps can be found in a similar axis that have apparently upright cristae. The fresh new paracone labial rib was salient, but rather than prior groove. There is absolutely no metacone labial rib. The fresh para-, meso-, and metastyles is salient. The lingual cingulum is extremely poor to absent including the entostyle.
The newest submitted design of the last lower premolar is typical to possess derived base pecoran and you may top pecoran within the which have a well-create mesolingual conid and you will posterior cristid , . More over, the presence of a tooth outlet into the p1 excludes a keen attribution in order to Prodremotherium otherwise Mosaicomeryx . Even when the pearly whites commonly since bunoselenodont as with Amphitragulus elegans, brand new referred specimens have quite primitive keeps, such a very shorter metastylid, a low top, a little postentocristid, and even though laterally compacted, bulged lingual cuspids towards down molars. All these qualities exclude an association to sometimes Dremotherium or Bedenomeryx , . Furthermore, the fresh new most small size regarding “Amphitragulus” quercyi are an identifying feature with the current Oligocene ruminants of European countries , , , .
Dremotherium cf. guthi regarding Gaimersheim1: step one kept maxillary bone on the enamel row M1-M3 (SNSB-BSPG 1952II4855). off Mailhat: dos left P4-M2 (MNHN LIM154). Babameryx engesseri off Gaimersheim1: step three kept fragmentary mandible which have m2 (SNSB-BSPG 1952II4818), (a) lingual, (b) labial, and you may (c) occlusal opinions. The scale bar was dos cm.
Holotype referred specimens pointers
Holotype. Jehenne (1987) has actually designated the new sample IPHEP LM.1969.MA.forty since the holotype. This might be a limited best maxillary limbs which have P3-M3 (, Fig. 2), away from Los angeles Milloque (MP29, France).
Known specimens. Mailhat MNHN LIM154 remaining P4-M2 (so it performs, Fig. six.2); Gaimersheim1 SNSB-BSPG 1952II4855 left maxillary that have M1-M3 (which functions, Fig. six.1); SNSB-BSPG 1952II4870 right m2.
Malfunction
Lower permanent dentition. The lower molar is quite improperly kept. Although not, so it larger ruminant and has now a totally selenomorph trigonid. The newest cristid try elongated as well as the cuspids are highest toward tooth quite pleated. The brand new metastylid are ripped.
Upper long lasting dentition. Brand new P4 is actually badly kept. All the lingual region try lost. The anterolabial cone and contains a failing labial rib. The newest prior stylid was broken. The posterior a person is quick but establish. No neocrista blocks the fresh new fossa. The fresh new quadratic upper molars in the occlusal view sustain a well-arranged metaconule some smaller compared to the latest protocone. New cusps are very selenomorph. This new preprotocrista and postmetaconulecrista try labially mainly based and you can some upright. The brand new postprotocrista is long, very curved, and labially based. It can be completely remote (SNSB-BSPG 1952II4855) or fused (MNHN LIM154) toward premetaconulecrista. The latter stops during the axis of your own mesostyle. The new posterior basin is 
keeps a little metaconule fold into the M2, large for the M3. The brand new paracone, possessing good paracone rib, plus the metacone, labially apartment and concave, was lined up. The new mesostyle and you will parastyle are setup. It function game small columns. The mesostyle are huge out-of M1 so you’re able to M3. New entostyle was weakened otherwise absent. Discover none good lingual cingulum close the brand new protocone neither an effective metastyle building a proper-setup cuspid. Yet not, short prior and posterior cingula exist.
Discussion
So it ruminant is relatively cutting-edge in-being better-selenodont and achieving no lingual cingula for the higher molars, usually present in Lophiomeryx, Gelocus, Iberomeryx, Mosaicomeryx, and you can Babameryx , , , [61–63], . As opposed to Bachitherium, which kinds lacks an anterior groove towards the large labial paracone rib , , . This new learned fossils much more similar to the state-of-the-art stem Pecora also to the fresh new ancient crown Pecora (such as for example Amphitragulus, Dremotherium, Bedenomeryx). The enamel is actually slightly pleated, whereas it is basically easy when you look at the Prodremotherium. Furthermore, the size of these pearly whites is a lot bigger than during the Prodremotherium on old Quercy stuff and Pech Desse . One’s teeth become more selenomorph compared to “Amphitragulus” and Bedenomeryx , , additionally the parastyle isn’t that anteriorly estimated than simply seen on the “Amphitragulus” feningrei , , . Studying the neodiagnosis proposed by Mennecart , these types of fossils were all of the proving options that come with Dremotherium. The fresh already discussed varieties in the Europe try D. feignouxi (sort of types) and D. guthi , , . Aforementioned try quite smaller compared to the sort varieties , similar to the fossils revealed here. D. feignouxi simply took place for the Agenian (MN1-MN2; ), whereas D. guthi stayed from inside the newest Oligocene (MP28-MP30; ). Although not, during the lack of a lot more well-preserved issue, a decisive assignation with the types is not possible.